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In foregut fermenters, bacterial cells are degraded later in the acidic stomach, where gastric lysozyme acts as a digestive enzyme (Ruiz et al., 1994) and bacterial biomass is the animal’s main source of protein (Stevens and Hume, 1995). The rumen harbors all three domains of life: bacteria (mostly Firmicutes and the Prevotella Bacteroides (Tajima et al., 2001a; Eckburg et al., 2005)); archaea (such as Methanobrevibacter and Thermoplasma (Tajima et al., 2001b)); and eukarya, both protozoa (ciliates and flagellates; (Orpin, 1976; Vogels et al., 1980) and fungi (anaerobic phycomycetes (Mackie, 1997)).Additional fermentation occurs in the large bowel and cecum of the cow, where the fermentable substrates are limited to the slower digesting polymers such as lignin and crystalline starches escaping foregut digestion and absorption, as well as some secreted mucins (Van Soest, 1994). Hindgut symbiotic bacteria continue the fermentation and also provide important vitamins for the host, such as vitamin K, thiamine and riboflavin (Burkholder and McVeigh, 1942; Nath and Meghal, 1961).
Results: Snus is manufactured and stored in a manner that causes it to deliver lower concentrations of some harmful chemicals than other tobacco products, although it can deliver high doses of nicotine. It is dependence forming, but does not appear to cause cancer or respiratory diseases. It may cause a slight increase in cardiovascular risks and is likely to be harmful to the unborn fetus, although these risks are lower than those caused by smoking.
